Meanwhile, an Oxford professor named Viviane Wembly is kidnapped by the Autobot Hot Rod. Both are brought to England and meet with Sir Edmund Burton, the last living member of the Witwiccan Order. Burton tells Cade and Viviane of the Transformers' origins, saying that the collision of Cybertron with Earth would spell certain doom. He tasks Viviane, the last descendant of Merlin, with obtaining his staff so they can prevent the impending apocalypse. Burton tells Yeager that, as a person defined by honor and dignity, he is the last knight. The talisman given to Yeager has wrapped around his arm by this point. The trio are forced to evacuate Burton's home when the MI6 and TRF arrives.
When he is gifted a Talisman by a dying Transformer knight, Cade Yeager finds himself pursued by both the Decepticons and the Transformers Reaction Force. He is soon recruited along with Viviane Wembly by Sir Edmund Burton, last of the Order of the Witwiccans, to once again save the world from a new impending threat.
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Cade's plane lands at Sir Burton's castle, where he and Bumblebee are shot at by a tank Transformer. Sir Burton apologizes and takes a look at the Talisman. Hot Rod arrives carrying Viviane, who is unhappy at being kidnapped. Once things calm down, Sir Burton explains he's the last of the Order of the Witwiccans. Cade's boorish behavior towards Viviane results in him being assaulted by Cogman. Sir Burton shows off portraits of his family tree and the Witwiccans, and relates how the Transformers have been helping humans for centuries, such as when Bumblebee and Hot Rod took part in World War II. They head to another building which houses the original Round Table used by King Arthur and his knights when they teamed up with the twelve Guardian Knights. Further there is a legend that a last knight would be found and thanks to possessing the Talisman, that's Cade. Only Merlin's DNA can unlock the power of the staff, which is where Viviane comes in as a direct descendant of Merlin. She has no idea where the staff might be though. Further discussion is halted as MI6 and TRF forces arrive to assault the castle. Despite one of the elderly Autobots attempting to defend the castle only to power down as a result of his aging, Hot Rod's time stopper weapon gives the group enough time to escape.
The chamber tumbles down until it comes to rest on another part of Cybertron. Inside, chaos reigns as the Autobots and Decepticons fight and Viviane grabs the staff. Optimus kicks Megatron through a wall shortly before another impact sends the chamber tumbling out of the sky. Bumblebee kills Nitro Zeus and blasts Quintessa as Optimus manages to distract her. Quintessa vanishes in a flash of blue light. Viviane finally pulls the staff free, shutting down the Ignition Chamber just before it crashes to the ground. Optimus dramatically drives out of the cloud of debris raised by its impact, carrying Cade and Viviane. As everyone reunites, Optimus gives another stirring speech about how Earth and Cybertron are now joined as one before the Autobots depart for Cybertron alongside the guardian knights.
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